The Holden Arboretum, Connect, Learn and Conserve

Holden Science and Research Staff

Mary A. Topa- Director of Science and Research

Plant physiology, physiological ecology, stress biology.

Steve Krebs - Director of the David G. Leach Research Station
Plant genetics and breeding, disease resistance, physiology of cold acclimation and winter freezing tolerance in rhododendron

Charlotte Chan- Research Associate

Don Whitney- David G. Leach Research Station Supervisor

David Burke - Assistant Scientist

Rhizosphere ecology, plant-microbe interactions, molecular microbial ecology, and plant ecology.

Kurt A. Smemo - Assistant Scientist

Soil biogeochemistry, ecosystem ecology, microbial ecology, global environmental change.

Beech-Maple forests of NE Ohio


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	*Holden Science and Research Staff* Mary A. Topa- Director of Science and Research *Plant physiology*, physiological ecology, stress biology. Steve Krebs - Director of the David G. Leach Research Station Plant genetics and breeding, disease resistance, physiology of cold acclimation and winter freezing tolerance in rhododendron Charlotte Chan- Research Associate Don Whitney- David G. Leach Research Station Supervisor David Burke - Assistant Scientist *Rhizosphere ecology*, plant-microbe interactions, molecular microbial ecology, and plant ecology. Kurt A. Smemo - Assistant Scientist *Soil biogeochemistry*, ecosystem ecology, microbial ecology, global environmental change.